. Tamura, Similarly, the sister-group relationship between Liza argentea and Aldrichetta had a higher posterior probability in the Bayesian analysis with Matrix 2 than Matrix 1. This suggests that a denser sampling of taxa, as in Matrix 2 relative to Matrix 1, improves the accuracy of phylogenetic inference (Hillis, 1996) Accordingly, the family was found to comprise 7 major clades that have radiated early from the ancestor to all current Mugilidae (Fig. 3). The phylogenetic relationships among Mugilidae genera, as inferred from the present analyses, are summarized in Fig. 1F. A detailed account of the phylogenetic placement of each Mugilidae genus is given below, where genera are listed in alphabetical order Agonostomus was paraphyletic with respect to Joturus (Fig. 3); A. monticola was phylogenetically closer to J. pichardi than both were to A. catalai. The nucleotide divergence between A. monticola and A. catalai at locus 16S, clade formed by Myxus elongatus and Neomyxus, with stronger support for the results inferred from Matrix 2 (Fig. 3) than Matrix 1, 2011.

. Cestraeus, represented by two species (C. goldiei and C. oxyrhinchus) was found to be monophyletic and a brother genus to Chaenomugil

L. Chelon-labrosus-grouped-with, L. Aurata, L. Ramada, L. Saliens, L. Richardsonii et al., dumerili and L. tricuspidens (Fig. 5A) to form a monophyletic subclade (Fig. 3), which turned out to exclusively comprise species distributed in Atlantic and Mediterranean waters or around southern Africa. The other Chelon species sampled, all from the Indo-Pacific, formed a distinct subclade together with Indo-Pacific Liza spp

D. Aurelle, R. Barthelemy, J. Quignard, M. Trabelsi, and E. Faure, Molecular Phylogeny of Mugilidae (Teleostei: Perciformes), The Open Marine Biology Journal, vol.2, issue.1, pp.29-37, 2008.
DOI : 10.2174/1874450800802010029

M. Autem and F. Bonhomme, El??ments de syst??matique biochimique chez les mugilid??s de M??diterran??e, Biochemical Systematics and Ecology, vol.8, issue.3, pp.305-308, 1980.
DOI : 10.1016/0305-1978(80)90065-4

J. C. Avise, Phylogeography: the history and formation of species, 2000.

H. Blel, N. Chatti, R. Besbes, S. Farjallah, A. Elouaer et al., Phylogenetic relationships in grey mullets (Mugilidae) in a Tunisian lagoon, Aquaculture Research, vol.25, issue.3, pp.268-275, 2008.
DOI : 10.1007/s10750-004-8772-5

J. C. Briggs, Fishes of Worldwide (Circumtropical) Distribution, Copeia, vol.1960, issue.3, pp.171-180, 1960.
DOI : 10.2307/1439652

F. Caldara, L. Bargelloni, L. Ostellari, E. Penzo, L. Colombo et al., Molecular Phylogeny of Grey Mullets Based on Mitochondrial DNA Sequence Analysis: Evidence of a Differential Rate of Evolution at the Intrafamily Level, Molecular Phylogenetics and Evolution, vol.6, issue.3, pp.416-424, 1996.
DOI : 10.1006/mpev.1996.0090

W. Chen, C. Bonillo, and G. Lecointre, Repeatability of clades as a criterion of reliability: a case study for molecular phylogeny of Acanthomorpha (Teleostei) with larger number of taxa, Molecular Phylogenetics and Evolution, vol.26, issue.2, pp.262-288, 2003.
DOI : 10.1016/S1055-7903(02)00371-8

W. Chen and R. L. Mayden, Molecular systematics of the Cyprinoidea (Teleostei: Cypriniformes), the world???s largest clade of freshwater fishes: Further evidence from six nuclear genes, Molecular Phylogenetics and Evolution, vol.52, issue.2, pp.544-549, 2009.
DOI : 10.1016/j.ympev.2009.01.006

W. Chen and R. L. Mayden, A Phylogenomic Perspective on the New Era of Ichthyology, BioScience, vol.60, issue.6, pp.421-432, 2010.
DOI : 10.1525/bio.2010.60.6.6

W. Chen, R. Ruiz-carus, and G. Orti, Relationships among four genera of mojarras (Teleostei: Perciformes: Gerreidae) from the western Atlantic and their tentative placement among percomorph fishes, Journal of Fish Biology, vol.36, issue.sb, pp.202-218, 2007.
DOI : 10.1007/BF00160154

D. Crosetti, J. C. Avise, F. Placidi, A. R. Rossi, and L. Sola, Geographic variability in the grey mullet Mugil cephalus: preliminary results of mtDNA and chromosome analyses, Aquaculture, vol.111, issue.1-4, pp.95-101, 1993.
DOI : 10.1016/0044-8486(93)90028-W

D. Crosetti, W. S. Nelson, and J. Avise, Pronounced genetic structure of mitochondrial DNA among populations of the circumglobally distributed grey mullet (Mugil cephalus), Journal of Fish Biology, vol.34, issue.1, pp.47-58, 1994.
DOI : 10.1007/BF00553878

R. C. Edgar, MUSCLE: multiple sequence alignment with high accuracy and high throughput, Nucleic Acids Research, vol.32, issue.5, pp.1792-1797, 2004.
DOI : 10.1093/nar/gkh340
URL : http://doi.org/10.1093/nar/gkh340

D. Erguden, M. Gurlek, D. Yaglioglu, and C. Turan, Genetic Identification and Taxonomic Relationship of Mediterranean Mugilid Species Based on Mitochondrial 16S rDNA Sequence Data, Journal of Animal and Veterinary Advances, vol.9, issue.2, pp.336-341, 2010.
DOI : 10.3923/javaa.2010.336.341

W. N. Eschmeyer and R. Fricke, Catalog of fishes, electronic version, 2011.

J. Felsenstein, Confidence Limits on Phylogenies: An Approach Using the Bootstrap, Evolution, vol.39, issue.4, pp.783-791, 1985.
DOI : 10.2307/2408678

E. Fraga, H. Schneider, M. Nirchio, E. Santa-brigida, L. F. Rodrigues-filho et al., Molecular phylogenetic analyses of mullets (Mugilidae, Mugiliformes) based on two mitochondrial genes, Journal of Applied Ichthyology, vol.41, issue.5, pp.598-604, 2007.
DOI : 10.1093/jhered/92.4.371

J. Ghasemzadeh, Phylogeny and Systematics of Indo-Pacific mullets (Teleostei: Mugilidae) with special reference to the mullets of Australia, Ph. D. dissertation. Macquarie University, 1998.

J. Ghasemzadeh, W. Ivantsoff, and . Aarn, Historical overview of mugilid systematics, with descriptions of Paramugil (Teleostei: Mugiliformes: Mugilidae), new genus, Aqua, J. Ichthyol. Aquat. Biol, vol.8, pp.9-22, 2004.

N. Goldman, Statistical tests of models of DNA substitution, Journal of Molecular Evolution, vol.46, issue.2, pp.182-198, 1993.
DOI : 10.1007/BF00166252

E. Gornung, P. Colangelo, and F. Annesi, 5S ribosomal RNA genes in six species of Mediterranean grey mullets: genomic organization and phylogenetic inference, Genome, vol.50, pp.787-795, 2007.

T. A. Hall, BIOEDIT: a user-friendly biological sequence alignement editor and analysis program for windows 95, NT. Nucl. Acids Symp. Ser, vol.98, issue.41, pp.95-98, 1999.

I. J. Harrison and G. J. Howes, The pharyngobranchial organ of mugilid fishes; its structure, variability, ontogeny, possible fonction and taxonomic utility, Bull. Brit. Mus. Nat. Hist., Zool, vol.57, pp.111-132, 1991.

I. J. Harrison, M. Nirchio, C. Oliveira, E. Ron, and J. Gaviria, A new species of mullet (Teleostei: Mugilidae) from Venezuela, with a discussion on the taxonomy of Mugil gaimardianus, Journal of Fish Biology, vol.41, issue.sa, pp.76-97, 2007.
DOI : 10.1007/s002270050312

I. J. Harrison and H. Senou, Mugilidae FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific, pp.2069-2108, 1999.

W. Hennig, Phylogenetic Systematics, Annual Review of Entomology, vol.10, issue.1, 1966.
DOI : 10.1146/annurev.en.10.010165.000525

S. Heras, M. Gonzalez-castro, and M. I. Roldan, Mugil curema in Argentinean waters: Combined morphological and molecular approach, Aquaculture, vol.261, issue.2, pp.473-478, 2006.
DOI : 10.1016/j.aquaculture.2006.07.003

S. Heras, M. I. Roldán, and M. Gonzalez-castro, Molecular phylogeny of Mugilidae fishes revised, Reviews in Fish Biology and Fisheries, vol.92, issue.2, pp.217-231, 2009.
DOI : 10.1007/s11160-008-9100-3

D. M. Hillis, Inferring complex phytogenies, Nature, vol.383, issue.6596, pp.130-131, 1996.
DOI : 10.1038/383130a0

D. M. Hillis, Taxonomic Sampling, Phylogenetic Accuracy, and Investigator Bias, Systematic Biology, vol.47, issue.1, pp.3-8, 1998.
DOI : 10.1080/106351598260987

J. P. Huelsenbeck and F. Ronquist, MRBAYES: Bayesian inference of phylogenetic trees, Bioinformatics, vol.17, issue.8, pp.754-755, 2001.
DOI : 10.1093/bioinformatics/17.8.754

A. Imsiridou, G. Minos, V. Katsares, N. Karaiskou, and A. Tsiora, Genetic identification and phylogenetic inferences in different Mugilidae species using 5S rDNA markers, Aquaculture Research, vol.96, issue.13, pp.1370-1379, 2007.
DOI : 10.1139/gen-44-5-893

B. W. Jamandre, J. Durand, and W. Tzeng, in the north-west Pacific as inferred from the mtDNA control region, Journal of Fish Biology, vol.60, issue.2, pp.393-407, 2009.
DOI : 10.1111/j.1095-8649.2009.02332.x

H. M. Ke, W. W. Lin, and H. W. Kao, ) in the Coastal Waters of Taiwan, Zoological Science, vol.26, issue.6, pp.421-428, 2009.
DOI : 10.2108/zsj.26.421.s1

A. G. Kluge, A concern for evidence and a phylogenetic hypothesis of relationships among Epicrates, 1989.

S. C. Lee, J. T. Chang, and Y. Y. Tsu, Genetic relationships of four Taiwan mullets (Pisces: Perciformes: Mugilidae), Journal of Fish Biology, vol.26, issue.1, pp.159-162, 1995.
DOI : 10.1577/1548-8659(1990)119<0002:GNFPLI>2.3.CO;2

B. Li, A. Dettaï, C. Cruaud, A. Couloux, M. Desoutter-meniger et al., RNF213, a new nuclear marker for acanthomorph phylogeny, Molecular Phylogenetics and Evolution, vol.50, issue.2, pp.345-363, 2009.
DOI : 10.1016/j.ympev.2008.11.013

C. Linnaeus, Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I, editio decima, reformata. Holmiae, 1758.

J. Liu, C. L. Brown, and T. Yang, Population genetic structure and historical demography of grey mullet, Mugil cephalus, along the coast of China, inferred by analysis of the mitochondrial control region, Biochemical Systematics and Ecology, vol.37, issue.5, pp.556-566, 2009.
DOI : 10.1016/j.bse.2009.09.002

J. Y. Liu, C. L. Brown, and T. B. Yang, Phylogenetic relationships of mullets (Mugilidae) in China Seas based on partial sequences of two mitochondrial genes, Biochemical Systematics and Ecology, vol.38, issue.4, pp.647-655, 2010.
DOI : 10.1016/j.bse.2010.01.007

S. Livi, L. Sola, and D. Crosetti, Phylogeographic relationships among worldwide populations of the cosmopolitan marine species, the striped gray mullet (Mugil cephalus), investigated by partial cytochrome b gene sequences, Biochemical Systematics and Ecology, vol.39, issue.2, pp.121-131, 2011.
DOI : 10.1016/j.bse.2011.01.006

G. Lüther, New characters for consideration in the taxonomy appraisal of grey mullets, J. Mar. Biol. Ass. India, vol.169, pp.1-9, 1982.

K. Mabuchi, M. Miya, Y. Azuma, and M. Nishida, Independent evolution of the specialized pharyngeal jaw apparatus in cichlid and labrid fishes, BMC Evolutionary Biology, vol.7, issue.1, p.10, 2007.
DOI : 10.1186/1471-2148-7-10

Y. Mandelik, T. Dayan, V. Chikatunov, and V. Kravchenko, Reliability of a Higher-Taxon Approach to Richness, Rarity, and Composition Assessments at the Local Scale, Conservation Biology, vol.50, issue.6, pp.1506-1515, 2007.
DOI : 10.1111/j.1523-1739.2007.00823.x

M. R. Menezes, M. Martin, and S. Naik, Interspecific genetic divergence in grey mullets from the Goa region, Aquaculture, vol.105, issue.2, pp.117-129, 1992.
DOI : 10.1016/0044-8486(92)90124-4

R. Murgia, G. Tola, S. N. Archer, S. Vallerga, and J. Hirano, Genetic Identification of Grey Mullet Species (Mugilidae) by Analysis of Mitochondrial DNA Sequence: Application to Identify the Origin of Processed Ovary Products (Bottarga), Marine Biotechnology, vol.4, issue.2, pp.119-126, 2002.
DOI : 10.1007/s10126-001-0074-6

J. S. Nelson, Fishes of the World, 2006.
DOI : 10.1002/9781119174844

M. Nirchio and H. Cequea, Karyology of Mugil liza and M. curema from Venezuela, Bol. Invest. Mar. Cost, vol.27, pp.45-50, 1998.

J. A. Nylander, MRMODELTEST v. 2. Program distributed by the author, Evolutionary Biology Centre, 2004.

S. Palumbi, A. Martin, S. Romano, W. O. Mcmillan, L. Stice et al., The simple fool's guide to PCR, version 2.0, p.47, 1991.

V. Papasotiropoulos, E. Klossa-kilia, S. Alahiotis, and G. Kilias, Molecular phylogeny of grey mullets (Teleostei: Mugilidae) in, 2007.

V. Papasotiropoulos, E. Klossa-kilia, G. Kilias, and S. Alahiotis, Genetic divergence and phylogenetic relationships in grey mullets (Teleostei: Mugilidae) using allozyme data, Biochemical Genetics, vol.39, issue.5/6, pp.155-168, 2001.
DOI : 10.1023/A:1010289224573

V. Papasotiropoulos, E. Klossa-kilia, G. Kilias, and S. Alahiotis, Genetic divergence and phylogenetic relationships in grey mullets (Teleostei: Mugilidae) based on PCR RFLP analysis of mtDNA segments, Biochemical Genetics, vol.40, issue.3/4, pp.71-86, 2002.
DOI : 10.1023/A:1015162304934

H. Philippe, A. Chenuil, and A. Adoutte, Can the Cambrian explosion be inferred through molecular phylogeny? Development, pp.15-25, 1994.

A. Rambaut, Se-Al: Sequence Alignment Editor ver. 2.0. (Program distributed by the author, 1996.

A. Rocha-olivares, N. M. Garber, and K. C. Stuck, High genetic diversity, large inter-oceanic divergence and historical demography of the striped mullet, Journal of Fish Biology, vol.26, issue.5, pp.1134-1149, 2000.
DOI : 10.1046/j.1365-294x.1999.00643.x

A. R. Rossi, M. Capula, D. Crosetti, D. E. Campton, and L. Sola, Genetic divergence and phylogenetic inferences in five species of Mugilidae (Pisces: Perciformes), Marine Biology, vol.131, issue.2, pp.213-218, 1998.
DOI : 10.1007/s002270050313

A. R. Rossi, M. Capula, D. Crosetti, L. Sola, and D. E. Campton, Allozyme variation in global populations of striped mullet, Mugil cephalus (Pisces: Mugilidae), Marine Biology, vol.131, issue.2, pp.203-212, 1998.
DOI : 10.1007/s002270050312

A. R. Rossi, A. Ungaro, S. De-innocentiis, D. Crosetti, and L. Sola, Phylogenetic Analysis of Mediterranean Mugilids by Allozymes and 16S mt-rRNA Genes Investigation: Are the Mediterranean Species of Liza Monophyletic?, Biochemical Genetics, vol.42, issue.9/10, pp.301-315, 2004.
DOI : 10.1023/B:BIGI.0000039806.12756.81

J. Sambrook, E. F. Fritschi, and T. Maniatis, Molecular cloning: a laboratory manual, 1989.

L. P. Schultz, A revision of the genera of mullets, fishes of the family Mugilidae, with descriptions of three new genera, Proc. U.S. Natl. Mus. 96, pp.377-395, 1946.
DOI : 10.5479/si.00963801.96-3204.377

L. P. Schultz, Family Mugilidea, Fishes of the Marshall and Marianas islands Families from Asymmetrontidae through Siganidae. Bull. U.S. Natl Mus, pp.310-322, 1953.

H. Senou, Phylogenetic interrelationships of the Mullets (Pisces: Mugilidae), 1988.

D. H. Setiamarga, M. Miya, Y. Yamanoue, K. Mabuchi, T. P. Satoh et al., Interrelationships of Atherinomorpha (medakas, flyingfishes, killifishes, silversides, and their relatives): The first evidence based on whole mitogenome sequences, Molecular Phylogenetics and Evolution, vol.49, issue.2, pp.598-605, 2008.
DOI : 10.1016/j.ympev.2008.08.008

R. G. Sevilla, A. Diez, M. Noren, O. Mouchel, M. Jerome et al., Primers and polymerase chain reaction conditions for DNA barcoding teleost fish based on the mitochondrial cytochrome b and nuclear rhodopsin genes, Molecular Ecology Notes, vol.13, issue.5, pp.730-734, 2007.
DOI : 10.1007/PL00006545

K. Shen, B. W. Jamandre, C. Hsu, W. Tzeng, and J. Durand, Plio-Pleistocene sea level and temperature fluctuations in the northwestern Pacific promoted speciation in the globally, 2011.

H. Shimodaira and M. Hasegawa, Multiple Comparisons of Log-Likelihoods with Applications to Phylogenetic Inference, Molecular Biology and Evolution, vol.16, issue.8, pp.1114-1116, 1999.
DOI : 10.1093/oxfordjournals.molbev.a026201

W. L. Smith and M. T. Craig, Casting the Percomorph Net Widely: The Importance of Broad Taxonomic Sampling in the Search for the Placement of Serranid and Percid Fishes, Copeia, vol.2007, issue.1, pp.35-55, 2007.
DOI : 10.1643/0045-8511(2007)7[35:CTPNWT]2.0.CO;2

W. L. Smith and W. C. Wheeler, Venom Evolution Widespread in Fishes: A Phylogenetic Road Map for the Bioprospecting of Piscine Venoms, Journal of Heredity, vol.97, issue.3, pp.206-217, 2006.
DOI : 10.1093/jhered/esj034

A. Stamatakis, RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models, Bioinformatics, vol.22, issue.21, pp.2688-2690, 2006.
DOI : 10.1093/bioinformatics/btl446

A. Stamatakis, P. Hoover, and J. Rougemont, A Rapid Bootstrap Algorithm for the RAxML Web Servers, Systematic Biology, vol.57, issue.5, pp.758-771, 2008.
DOI : 10.1080/10635150802429642

M. L. Stiassny, What are grey mullets?, Bull. Mar. Sci, vol.52, pp.197-219, 1993.

D. L. Swofford, PAUP*. Phylogenetic analysis using parsimony (*and other methods), version 4, 2002.

K. Tamura and M. Nei, Estimation of the number of nucleotide substitutions in the control region of mitochondrial DNA in humans and chimpanzees, Mol. Biol. Evol, vol.10, pp.512-526, 1993.

K. Tamura, D. Peterson, N. Peterson, G. Stecher, M. Nei et al., MEGA5: Molecular Evolutionary Genetics Analysis Using Maximum Likelihood, Evolutionary Distance, and Maximum Parsimony Methods, Molecular Biology and Evolution, vol.28, issue.10, pp.2731-2739, 2011.
DOI : 10.1093/molbev/msr121

J. M. Thomson, The Mugilidae of Australia and Adjacent Seas, Marine and Freshwater Research, vol.5, issue.1, pp.70-131, 1954.
DOI : 10.1071/MF9540070

J. M. Thomson, The grey mullets. Oceanogr, Mar. Biol. Annu. Rev, vol.4, pp.301-355, 1966.

J. M. Thomson, The Mugilidae of the world, Mem. Queensl. Mus, vol.43, pp.457-562, 1997.

C. Turan, M. Caliskan, and H. Kucuktas, Phylogenetic relationships of nine mullet species (Mugilidae) in the Mediterranean Sea, Hydrobiologia, vol.30, issue.1-3, pp.45-51, 2005.
DOI : 10.1007/s10750-004-8772-5

J. L. Villaseñor, G. Ibarra-manriquez, J. A. Meave, and E. Ortiz, Higher Taxa as Surrogates of Plant Biodiversity in a Megadiverse Country, Conservation Biology, vol.28, issue.1, pp.232-238, 2005.
DOI : 10.1046/j.1523-1739.1996.10010155.x

R. D. Ward, T. S. Zemlak, B. H. Innes, P. R. Last, and P. D. Hebert, DNA barcoding Australia's fish species, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.11, issue.6897, pp.1847-1857, 2005.
DOI : 10.1093/molbev/msg133

J. J. Wiens and M. C. Morrill, Missing Data in Phylogenetic Analysis: Reconciling Results from Simulations and Empirical Data, Systematic Biology, vol.60, issue.5, pp.719-731, 2011.
DOI : 10.1093/sysbio/syr025

M. Cephalus, Chile) 375. M. cephalus (Chile) A 0

. Moolgarda-sp, Reunion) 221a. Moolgarda seheli (Tanzania)

. Moolgarda-sp, Reunion) 241. Valamugil sp. (Taiwan) 238. Valamugil sp. (Fiji) 239. Valamugil sp