B. J. Morgan, J. R. Tillett, and . Ovenden, A review of the application of molecular genetics for fisheries management and conservation of sharks and rays, J. Fish Biol, vol.80, pp.1789-1843, 2012.

W. T. White and P. R. Last, A review of the taxonomy of chondrichthyan fishes: a modern perspective, Journal of Fish Biology, vol.59, issue.5, pp.901-917, 2012.
DOI : 10.1111/j.1095-8649.2011.03192.x

W. T. White, S. J. Blaber, and J. F. Craig, The current status of elasmobranchs: biology, fisheries and conservation, Journal of Fish Biology, vol.80, issue.5, pp.897-900, 2012.
DOI : 10.1111/j.1095-8649.2012.03268.x

P. R. Last, W. T. White, and J. , Descriptions of new Australian Chondrichthyans, CSIRO Mar, Atm. Res, vol.358, 2008.

W. T. White and P. M. Kyne, The status of chondrichthyan conservation in the Indo-Australasian region, Journal of Fish Biology, vol.91, issue.9, pp.2090-2117, 2010.
DOI : 10.1111/j.1095-8649.2010.02654.x

S. J. Blaber, C. M. Dichmont, W. White, R. Buckworth, L. Sadiyah et al., Elasmobranchs in southern Indonesian fisheries: the fisheries, the status of the stocks and management options, Elasmobranchs in southern Indonesian fisheries: the fisheries, the status of the stocks and management options, pp.367-391, 2009.
DOI : 10.1007/s11160-009-9110-9

V. Y. Lam and Y. Sadovy-de-mitcheson, The sharks of South East Asia - unknown, unmonitored and unmanaged, Fish and Fisheries, vol.23, issue.1, pp.51-74, 2011.
DOI : 10.1111/j.1467-2979.2010.00383.x

N. C. Aschliman, M. Nishida, M. Miya, J. G. Inoue, K. M. Rosana et al., Body plan convergence in the evolution of skates and rays (Chondrichthyes: Batoidea), Molecular Phylogenetics and Evolution, vol.63, issue.1, pp.63-91, 2012.
DOI : 10.1016/j.ympev.2011.12.012

D. Armitage, Socio-institutional dynamics and the political ecology of mangrove forest conservation in Central Sulawesi, Indonesia, Global Environmental Change, vol.12, issue.3, pp.203-217, 2002.
DOI : 10.1016/S0959-3780(02)00023-7

J. F. Bruno and E. R. Selig, Regional Decline of Coral Cover in the Indo-Pacific: Timing, Extent, and Subregional Comparisons, PLoS ONE, vol.360, issue.8, p.711, 2007.
DOI : 10.1371/journal.pone.0000711.s010

M. Heazle and J. G. Butcher, Fisheries depletion and the state in Indonesia: Towards a regional regulatory regime, Marine Policy, vol.31, issue.3, pp.276-286, 2007.
DOI : 10.1016/j.marpol.2006.08.006

C. H. Ainsworth, T. J. Pitcher, and C. Rotinsulu, Evidence of fishery depletions and shifting cognitive baselines in Eastern Indonesia, Biological Conservation, vol.141, issue.3, pp.848-859, 2008.
DOI : 10.1016/j.biocon.2008.01.006

B. M. Manjaji-matsumoto and P. R. Last, Himantura leoparda sp. nov., a new whipray (Myliobatoidei: Dasyatidae) from the Indo?Pacific, Descriptions of new Australian Chondrichthyans, CSIRO Mar. Atm. Res. Pap. 22. CSIRO, pp.293-301, 2008.

P. R. Last and J. D. Stevens, Sharks and rays of Australia, 1994.

P. R. Last, L. J. Compagno, and D. , FAO species identification guide for fishery purposes, The living marine resources of the Western Central Pacific Batoid fishes, chimaeras and bony fishes, pp.1479-1505, 1999.

N. Annandale, Report on the fishes taken by the Bengal fisheries steamer 'Golden Crown', Part I, Batoidei, Mem, Indian Mus, vol.2, pp.1-60, 1909.

P. Bleeker, Bijdrage tot de kennis der Plagiostomen van den Indischen archipel, Verhandelingen van het, Bataviaasch Genootschap van Kunsten en Wettenschappen, vol.24, pp.1-92, 1852.

E. Mayr, Systematics and the origin of species from the viewpoint of a zoologist, 1942.

R. D. Ward, R. Hanner, and P. D. Hebert, The campaign to DNA barcode all fishes, FISH-BOL, Journal of Fish Biology, vol.57, issue.Suppl. B, pp.329-356, 2009.
DOI : 10.1111/j.1095-8649.2008.02080.x

F. Cerutti-pereyra, M. G. Meekan, N. V. Wei, O. O. Shea, C. J. Bradshaw et al., Identification of Rays through DNA Barcoding: An Application for Ecologists, PLoS ONE, vol.23, issue.6, p.36479, 2012.
DOI : 10.1371/journal.pone.0036479.s003

S. Ratnasingham and P. D. Hebert, BARCODING: bold: The Barcode of Life Data System (http://www.barcodinglife.org), Molecular Ecology Notes, vol.360, issue.3, pp.355-364, 2007.
DOI : 10.1111/j.1471-8286.2007.01678.x

S. R. Palumbi, A. P. Martin, S. Romano, W. O. Mcmillan, L. Stice et al., The Simple Fool's Guide to PCR, Version 2.0, p.46, 1991.

T. Kitamura, A. Takemura, S. Watabe, T. Taniuchi, and M. Shimizu, Molecular phylogeny of the sharks and rays of Superorder Squalea based on mitochondrial cytochrome b gene, Fisheries Sci, pp.62-340, 1996.

K. Tamura, Estimation of the number of nucleotide substitutions when there are strong transitiontransversion and G+C content biases, Mol. Biol. Evol, vol.9, pp.678-687, 1992.

K. Tamura, D. Peterson, N. Peterson, G. Stecher, M. Nei et al., MEGA 5: Molecular evolutionary genetics analysis using maximum likelihood, evolutionary distance, and maximum parsimony methods, Mol. Biol. Evol, pp.28-2731, 2011.
DOI : 10.1093/molbev/msr121

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3203626

T. A. Hall, BIOEDIT: a user-friendly biological sequence alignement editor and analysis program for Windows 95, NT Nucl. Acids Symp. Ser, vol.98, pp.41-95, 1999.

S. Lê, J. Josse, and F. Husson, FACTOMINER: an R package for multivariate analysis, J. Stat. Software, vol.25, issue.1, 2008.

R. Ihaka and R. Gentleman, R: a language for data analysis and graphics, J. Comput. Graph. Stat, vol.5, pp.299-314, 1996.

D. Tautz, P. Arctander, A. Minelli, R. H. Thomas, and A. P. Vogler, A plea for DNA taxonomy, Trends in Ecology & Evolution, vol.18, issue.2, pp.70-74, 2003.
DOI : 10.1016/S0169-5347(02)00041-1

R. D. Ward, T. S. Zemlak, B. H. Innes, P. R. Last, and P. D. Hebert, DNA barcoding Australia's fish species, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.11, issue.6897, pp.1847-1857, 2005.
DOI : 10.1093/molbev/msg133

A. Valentini, F. Pompanon, and P. Taberlet, DNA barcoding for ecologists, Trends in Ecology & Evolution, vol.24, issue.2, pp.110-117, 2008.
DOI : 10.1016/j.tree.2008.09.011

URL : https://hal.archives-ouvertes.fr/halsde-00378005

L. G. Cook, R. D. Edwards, M. D. Crisp, and N. B. Hardy, Need morphology always be required for new species descriptions?, Invertebrate Systematics, vol.24, issue.3, pp.322-326, 2010.
DOI : 10.1071/IS10011

G. J. Naylor, J. N. Caira, K. Jensen, K. A. Rosana, W. T. White et al., A DNA Sequence???Based Approach To the Identification of Shark and Ray Species and Its Implications for Global Elasmobranch Diversity and Parasitology, Bulletin of the American Museum of Natural History, vol.367, pp.367-368, 2012.
DOI : 10.1206/754.1

B. M. Manjaji, Taxonomy and phylogenetic systematics of the Indo-Pacific whip-tailed stingray genus Himantura Müller and Henle 1837 (Chondrichthyes: Myliobatiformes: Dasyatidae), 2004.

P. Borsa, I. S. Arlyza, M. Laporte, and P. Berrebi, Population genetic structure of blue-spotted maskray Neotrygon kuhlii and two other Indo-West Pacific stingray species (Myliobatiformes: Dasyatidae), inferred from size-polymorphic intron markers, Journal of Experimental Marine Biology and Ecology, vol.438, pp.438-470, 2012.
DOI : 10.1016/j.jembe.2012.09.010

URL : https://hal.archives-ouvertes.fr/ird-00747306

R. D. Ward, B. H. Holmes, W. T. White, and P. R. Last, DNA barcoding Australian chondrichthyans: results and potential uses in conservation, Mar. Freshw. Res, pp.59-57, 2008.
DOI : 10.1071/mf07148

URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.509.6954

M. Adrim, Studi biodiversitas hiu dan pari di Indonesia, Laporan proyek kompetitif, LIPI-P2O, p.59, 2008.

. Fig and . Arlyza, Himantura uarnak species complex Pictures of specimens analyzed in the present study, with details on sex, disk width (DW), and multiple-locus nuclear genotype [Cluster 1, Cluster 3, or Cluster 4 according to Sampling locations (fish landing sites) and dates were, 2007.

B. Selayar and . Sea, IR 102); SingarajaIR 112); Pemba Island, 2009.

E. C. Taiwan and . Sea, wjc 637); north-west of Weipa, Gulf of Carpentaria, CSIRO H2903.01)., IR 036 (male, 1070 mm DW, cluster 4) IR 037 (female, 1180 mm DW, presumed F1cluster 1 x cluster 3) IR 038 (female, 660 mm DW, 1991.

. S. Fig, (continued) IR 055 (male, 1130 mm DW, cluster 1) IR 056 (female, 1070 mm DW, cluster 1) IR 057 (female, 1360 mm DW

. S. Fig, (continued) IR 064 (female, 1430 mm DW, cluster 4) IR 065 (male, 1090 mm DW, cluster 4) IR 066 (female, 940 mm DW

. S. Fig, (continued) IR 073 (male, 1160 mm DW, cluster 1) IR 074 (female, 1220 mm DW, cluster 4) IR 075 (female, 1300 mm DW

. S. Fig, (continued) IR 082 (female, 1210 mm DW, cluster 1) IR 083 (female, 1020 mm DW, cluster 1) IR 084 (female, 1200 mm DW