W. W. Adams, C. R. Zarter, K. E. Mueh, V. Amiard, and B. Demmig-adams, Energy Dissipation and Photoinhibition: A Continuum of Photoprotection, Gene Regulation, and Environment. Advances in Photosynthesis and Respiration, vol.22, pp.49-64, 2006.
DOI : 10.1007/1-4020-3579-9_5

G. S. Alves, L. F. Torres, E. Déchamp, J. Breitler, T. Joët et al., Differential fine-tuning of gene expression regulation in coffee leaves by CcDREB1D promoter haplotypes under water deficit, Journal of Experimental Botany, vol.68, issue.11
DOI : 10.1093/jxb/erx166

W. L. Araujo, P. C. Dias, G. A. Moraes, E. F. Celin, R. L. Cunha et al., Limitations to photosynthesis in coffee leaves from different canopy positions, Plant Physiology and Biochemistry, vol.46, issue.10, pp.884-890, 2008.
DOI : 10.1016/j.plaphy.2008.05.005

P. A. Armond, O. Bjorkman, and L. A. Staehelin, Dissociation of supramolecular complexes in chloroplast membranes A manifestation of heat damage to the photosynthetic apparatus, Biochimica et Biophysica Acta (BBA) - Biomembranes, vol.601, pp.433-442, 1980.
DOI : 10.1016/0005-2736(80)90547-7

J. Azcon-bieto, Inhibition of Photosynthesis by Carbohydrates in Wheat Leaves, PLANT PHYSIOLOGY, vol.73, issue.3, pp.681-686, 1983.
DOI : 10.1104/pp.73.3.681

J. Azcon-bieto, H. Lambers, and D. A. Day, Effect of photosynthesis and carbohydrate status on respiratory rates and the involvement of the alternative path in leaf respiration, Plant Physiol, vol.723, pp.598-603, 1983.

K. D. Batista, W. L. Araújo, W. C. Antunes, P. C. Cavatte, G. A. Moraes et al., Photosynthetic limitations in coffee plants are chiefly governed by diffusive factors, Trees, vol.128, issue.2, pp.459-468, 2012.
DOI : 10.1093/treephys/25.6.753

J. C. Begara-morales, B. Sánchez-calvo, M. Chaki, C. Mata-pérez, R. Valderrama et al., -nitrosylation, Journal of Experimental Botany, vol.66, issue.19, pp.5983-5996, 2015.
DOI : 10.1093/jxb/erv306

L. P. Bidel, G. Chomicki, F. Bonini, L. Mondolot, J. Soulé et al., Dynamics of flavonol accumulation in leaf tissues under different UV-B regimes in Centella asiatica (Apiaceae), Planta, vol.53, issue.4, pp.545-559, 2015.
DOI : 10.1093/pcp/pcs088

URL : https://hal.archives-ouvertes.fr/hal-01269253

L. P. Bidel, S. Meyer, Y. Goulas, Y. Cadot, C. et al., Responses of epidermal phenolic compounds to light acclimation: In vivo qualitative and quantitative assessment using chlorophyll fluorescence excitation spectra in leaves of three woody species, Journal of Photochemistry and Photobiology B: Biology, vol.88, issue.2-3, pp.163-179, 2007.
DOI : 10.1016/j.jphotobiol.2007.06.002

M. M. Bradford, A rapid and sensitive method for the quantitation of microgram quantities of protein utilizing the principle of protein-dye binding, Analytical Biochemistry, vol.72, issue.1-2, pp.248-254, 1976.
DOI : 10.1016/0003-2697(76)90527-3

J. G. Buwalda and G. Noga, Intra???plant differences in leaf chlorophyll fluorescence parameters in perennial fruiting plants, New Zealand Journal of Crop and Horticultural Science, vol.4, issue.4, pp.373-380, 1994.
DOI : 10.1007/BF00240982

URL : http://www.tandfonline.com/doi/pdf/10.1080/01140671.1994.9513848?needAccess=true

I. Cakmak and H. Marschner, Magnesium Deficiency and High Light Intensity Enhance Activities of Superoxide Dismutase, Ascorbate Peroxidase, and Glutathione Reductase in Bean Leaves, PLANT PHYSIOLOGY, vol.98, issue.4, pp.1222-1227, 1992.
DOI : 10.1104/pp.98.4.1222

C. Campa, L. Mondolot, A. Rakotondravao, L. P. Bidel, A. Gargadennec et al., A survey of mangiferin and hydroxycinnamic acid ester accumulation in coffee (Coffea) leaves: biological implications and uses, Annals of Botany, vol.110, issue.3, pp.595-613, 2008.
DOI : 10.1093/aob/mcs119

URL : https://hal.archives-ouvertes.fr/hal-01267806

D. C. Close and C. Mcarthur, Rethinking the role of many plant phenolics -protection from photodamage not herbivores? Oikos 99, 2002.

I. Couée, C. Sulmon, G. Gouesbet, E. Amrani, and A. , Involvement of soluble sugars in reactive oxygen species balance and responses to oxidative stress in plants, Journal of Experimental Botany, vol.57, issue.3, pp.449-459, 2006.
DOI : 10.1093/jxb/erj027

D. Matta and F. M. , Ecophysiological constraints on the production of shaded and unshaded coffee: a review. Field Crops Res, pp.99-114, 2004.

T. A. Day, Relating UV-B radiation screening effectiveness of foliage to absorbing-compound concentration and anatomical characteristics in a diverse group of plants, Oecologia, vol.335, issue.4, pp.542-550, 1993.
DOI : 10.1038/335154a0

S. Dussert, M. W. Davey, A. Laffargue, S. Doulbeau, R. Swennen et al., Oxidative stress, phospholipid loss and lipid hydrolysis during drying and storage of intermediate seeds, Physiologia Plantarum, vol.1240, issue.2, pp.192-204, 2006.
DOI : 10.1074/jbc.M205375200

W. R. Edwards, J. A. Hall, A. R. Rowlan, T. Schneider-barfield, T. J. Sun et al., Light filtering by epidermal flavonoids during the resistant response of cotton to Xanthomonas protects leaf tissue from light-dependent phytoalexin toxicity, Phytochemistry, vol.69, issue.12, pp.2320-2328, 2008.
DOI : 10.1016/j.phytochem.2008.05.021

J. I. Fahl, M. L. Carelli, J. Vega, and A. C. Magalhaes, L.), Journal of Horticultural Science, vol.69, issue.1, pp.161-169, 1994.
DOI : 10.1007/978-3-642-68090-8_4

B. R. Fondy and D. R. Geiger, Diurnal Pattern of Translocation and Carbohydrate Metabolism in Source Leaves of Beta vulgaris L., PLANT PHYSIOLOGY, vol.70, issue.3, pp.671-676, 1982.
DOI : 10.1104/pp.70.3.671

C. H. Foyer and B. Halliwell, The presence of glutathione and glutathione reductase in chloroplasts: A proposed role in ascorbic acid metabolism, Planta, vol.26, issue.1, pp.21-25, 1976.
DOI : 10.1007/BF00386001

S. C. Grace, L. , and B. A. , Energy dissipation and radical scavenging by the plant phenylpropanoid pathway, Philosophical Transactions of the Royal Society B: Biological Sciences, vol.355, issue.1402, pp.1499-1510, 2000.
DOI : 10.1098/rstb.2000.0710

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1692864

S. C. Grace, B. A. Logan, A. , and W. W. Iii, Seasonal differences in foliar content of chlorogenic acid, a phenylpropanoid antioxidant, in Mahonia repens, Plant, Cell and Environment, vol.45, issue.5, 1998.
DOI : 10.1006/abbi.1996.0331

R. E. Hausler, L. Heinrichs, J. Scmitz, and U. Flugge, How Sugars Might Coordinate Chloroplast and Nuclear Gene Expression during Acclimation to High Light Intensities, Molecular Plant, vol.7, issue.7, pp.1121-1137, 2014.
DOI : 10.1093/mp/ssu064

J. X. He, J. Wang, and H. G. Liang, Effects of water stress on photochemical function and protein metabolism of photosystem II in wheat leaves, Physiologia Plantarum, vol.56, issue.4, pp.771-777, 1995.
DOI : 10.1007/978-94-009-0511-5_47

S. Henkes, U. Sonnewald, R. Badur, R. Flachmann, and M. Stitt, A Small Decrease of Plastid Transketolase Activity in Antisense Tobacco Transformants Has Dramatic Effects on Photosynthesis and Phenylpropanoid Metabolism, THE PLANT CELL ONLINE, vol.13, issue.3, pp.535-551, 2001.
DOI : 10.1105/tpc.13.3.535

I. Hernandez, V. Breusegem, and F. , Opinion on the possible role of flavonoids as energy escape valves: Novel tools for nature's Swiss army knife?, Plant Science, vol.179, issue.4, 2010.
DOI : 10.1016/j.plantsci.2010.06.001

M. S. Islam, M. Yoshimoto, K. Ishiguro, S. Okuno, Y. et al., Effect of artificial shading and temperature on radical scavenging activity and polyphenolic composition in sweepotato (Ipomea batatas L.) leaves, J. Am. Soc. Hort. Sci, vol.128, pp.182-187, 2003.

M. A. Jansen, Ultraviolet-B radiation effects on plants: induction of morphogenic responses, Physiologia Plantarum, vol.16, issue.3, pp.423-429, 2002.
DOI : 10.1104/pp.102.3.783

P. Klein, S. Murray, W. Solano, C. Montagnon, T. Schilling et al., An extensive study on the genetic diversity of Arabica species evidences two genetic groups in Ethiopia and a sub-group in Yemen, Proceedings of the 26th International Conference on Coffee Science, pp.13-19, 2016.

C. Lu, J. Zhang, Q. Zhang, L. Li, and T. Kuang, Modification of photosystem II photochemistry in nitrogen deficient maize and wheat plants, Journal of Plant Physiology, vol.158, issue.11, pp.1423-1430, 2001.
DOI : 10.1078/0176-1617-00501

E. A. Maher, N. J. Bate, W. Ni, Y. Elkin, R. A. Dixon et al., Increased disease susceptibility of transgenic tobacco plants with suppressed levels of performed phenylpropanoid products, Proc. Nat. Acad. Sci. U.S.A. 91, pp.7802-7806, 1994.

V. Mahesh, R. Million-rousseau, P. Ullmann, N. Chabrillange, J. Bustamante et al., Functional characterization of two p-coumaroyl ester 3???-hydroxylase genes from coffee tree: evidence of a candidate for chlorogenic acid biosynthesis, Plant Molecular Biology, vol.247, issue.1-2, pp.145-159, 2007.
DOI : 10.1093/oxfordjournals.pcp.a029422

URL : https://hal.archives-ouvertes.fr/hal-00145089

S. C. Martins, W. L. Araujo, T. Tohge, A. R. Fernie, and F. M. Damatta, In high-ligh-acclimated coffee plants the metabolic machinery is adjusted to avoid oxidative stress rather than to benefit from extra light enhancement in photosynthetic yield, PLoS ONE, vol.9, 2014.

S. C. Martins, J. Galmés, P. C. Cavatte, L. F. Pereira, M. C. Ventrella et al., Understanding the Low Photosynthetic Rates of Sun and Shade Coffee Leaves: Bridging the Gap on the Relative Roles of Hydraulic, Diffusive and Biochemical Constraints to Photosynthesis, PLoS ONE, vol.105, issue.194, 2014.
DOI : 10.1371/journal.pone.0095571.s003

S. Mole, J. A. Ross, and P. G. Waterman, Light-induced variation in phenolic levels in foliage of rainforest plants, I. Chemical changes Histolocalisation de substances phénoliques intervenant lors d'interactions plante-pathogène chez le Tournesol et la Vigne, J. Chem. Ecol. Acta Bot. Gallica, vol.14, issue.144, pp.1-21, 1080.

R. Müller-xing, Q. Xing, and J. Goodrich, Footprints of the sun: memory of UV and light stress in plants, Front. Plant Sci, vol.5, p.474, 2014.

Y. Munekage, M. Hashimoto, C. Miyake, K. Tomizawa, T. Endo et al., Cyclic electron flow around photosystem I is essential for photosynthesis, Nature, vol.43, issue.6991, pp.579-582, 1038.
DOI : 10.1023/A:1020459416987

D. A. Myers, R. B. Thomas, and E. H. Delucia, Photosynthetic responses of loblolly pine (Pinus taeda) needles to experimental reduction in sink demand, Tree Physiology, vol.19, issue.4-5, pp.235-242, 1999.
DOI : 10.1093/treephys/19.4-5.235

Y. Nakano and K. Asada, Hydrogen peroxide as scavenged by ascorbatespecific peroxidase in spinach chloroplasts, Plant Cell Physiol, vol.2, pp.867-880, 1981.

A. T. Netto, E. Campostrini, J. Gonc, J. G. De-oliveira, and R. E. Bressan-smith, Photosynthetic pigments, nitrogen, chlorophyll a fluorescence and SPAD-502 readings in coffee leaves, Scientia Horticulturae, vol.104, issue.2, pp.199-209, 2005.
DOI : 10.1016/j.scienta.2004.08.013

Q. T. Nguyen, T. Kozai, G. Niu, and U. V. Nguyen, Photosynthetic characteristics of coffee (Coffea arabusta) plantlets in vitro in response to different CO2 concentrations and light intensities, Plant Cell, Tissue and Organ Culture, vol.55, issue.2, pp.133-139, 1998.
DOI : 10.1023/A:1006224026639

M. A. Nunes, J. C. Ramalho, and M. A. Dias, Effect of Nitrogen Supply on the Photosynthetic Performance of Leaves from Coffee Plants Exposed to Bright Light, Journal of Experimental Botany, vol.44, issue.5, pp.893-899, 1993.
DOI : 10.1093/jxb/44.5.893

M. J. Paul and C. H. Foyer, Sink regulation of photosynthesis, Journal of Experimental Botany, vol.52, issue.360, 2001.
DOI : 10.1093/jexbot/52.360.1383

URL : https://academic.oup.com/jxb/article-pdf/52/360/1383/18026874/521383.pdf

M. J. Paul and T. K. Pellny, Carbon metabolite feedback regulation of leaf photosynthesis and development, Journal of Experimental Botany, vol.54, issue.382, pp.539-547, 2003.
DOI : 10.1093/jxb/erg052

J. C. Ramalho, T. L. Pons, H. W. Groenveld, H. G. Azinheira, and M. A. Nunes, Photosynthetic acclimation to high light conditions in mature leaves of Coffea arabica L: role of xanthopylls, quenching mechanisms and nitrogen nutrition. Austr, J. Plant Physiol, vol.27, pp.43-51, 2000.

J. C. Ramalho, T. L. Pons, H. W. Groeneveld, and M. A. Nunes, Photosynthetic responses of Coffea arabica leaves to a short-term high light exposure in relation to N availability, Physiologia Plantarum, vol.83, issue.1, pp.229-239, 1997.
DOI : 10.1007/BF00035461

A. Rani, S. Kumar-vats, M. Sharma, and S. Kumar, Catechin promotes growth of Arabidopsis thaliana with concomitant changes in vascular system, photosynthesis and hormone content, Biologia Plantarum, vol.68, issue.4, pp.779-782, 2011.
DOI : 10.1021/jf9807263

T. M. Robson, K. Klem, O. Urban, and M. A. Jansen, Re-interpreting plant morphological responses to UV-B radiation, Plant, Cell & Environment, vol.200, issue.5, pp.856-866, 2015.
DOI : 10.1038/200761a0

J. S. Roden and M. C. Ball, The Effect of Elevated [CO2] on Growth and Photosynthesis of Two Eucalyptus Species Exposed to High Temperatures and Water Deficits, Plant Physiology, vol.111, issue.3, pp.909-919, 1996.
DOI : 10.1104/pp.111.3.909

M. Stitt, A. Von-schaewen, and L. Willmitzer, ???Sink??? regulation of photosynthetic metabolism in transgenic tobacco plants expressing yeast invertase in their cell wall involves a decrease of the Calvin-cycle enzymes and an increase of glycolytic enzymes, Planta, vol.183, issue.1, pp.40-50, 1007.
DOI : 10.1007/BF00197565

B. J. Strasser and R. J. Strasser, Measuring Fast Fluorescence Transients to Address Environmental Questions: The JIP-Test, Proceedings of the Xth International Photosynthesis Congress, 1995.
DOI : 10.1007/978-94-009-0173-5_1142

R. J. Strasser, A. Srivastava, and M. Tsimilli-michael, The fluorescence transient as a tool to characterize and screen photosynthetic samples, Probing Photosynthesis: Mechanisms, Regulation and Adaptation, pp.445-483, 2000.

A. J. Strauss, G. H. Kruger, R. J. Strasser, V. Heerden, and P. D. , Ranking of dark chilling tolerance in soybean genotypes probed by the chlorophyll a fluorescence transient O-J-I-P, Environmental and Experimental Botany, vol.56, issue.2, pp.147-157, 2006.
DOI : 10.1016/j.envexpbot.2005.01.011

C. Sundby, A. Melis, P. Maenpaa, and B. Andersson, Temperature-dependent changes in the antenna size of Photosystem II. Reversible conversion of Photosystem II?? to Photosystem II??, Biochimica et Biophysica Acta (BBA) - Bioenergetics, vol.851, issue.3, pp.475-483, 1986.
DOI : 10.1016/0005-2728(86)90084-8

U. Takahama and T. Oniki, A peroxidase/phenolics/ascorbate system can scavenge hydrogen peroxide in plant cells, Physiologia Plantarum, vol.264, issue.4, pp.845-852, 1997.
DOI : 10.1016/S1360-1385(97)82729-X

R. Tegelberg, R. Julkunen-tiitto, and P. J. Aphalo, Red : far-red light ratio and UV-B radiation: their effects on leaf phenolics and growth of silver birch seedlings, Plant, Cell and Environment, vol.200, issue.8, pp.1005-1013, 2004.
DOI : 10.1046/j.1365-313x.2001.01001.x

E. Thioune, J. Mccarthy, T. Gallagher, O. , B. R. et al., A humidity shock leads to rapid, temperature dependent changes in coffee leaf physiology and gene expression The role of the hexose transporter in the chloroplasts of Arabidopsis thaliana L, Tree Physiol. Planta, vol.37, issue.195, pp.1-13, 1007.

L. Urban, A. , and L. , Girdling decreases photosynthetic electron fluxes and induces sustained photoprotection in mango leaves, Tree Physiology, vol.27, issue.3, pp.345-352, 2007.
DOI : 10.1093/treephys/27.3.345

URL : https://academic.oup.com/treephys/article-pdf/27/3/345/4690560/27-3-345.pdf

L. Urban, M. Léchaudel, L. , P. S. Zimmermann, P. Rombauts et al., Effect of fruit load and girdling on leaf photosynthesis in Mangifera indica L Genome-wide analysis of hydrogen peroxide-regulated gene expression in Arabidopsis reveals a high light-induced transcriptional cluster involved in anthocyanin biosynthesis, J. Exp. Bot. Plant Physiol, vol.55, issue.139, pp.2075-2085, 1104.

R. Villar, A. A. Held, and J. Merino, Dark Leaf Respiration in Light and Darkness of an Evergreen and a Deciduous Plant Species, Plant Physiology, vol.107, issue.2, pp.421-427, 1995.
DOI : 10.1104/pp.107.2.421

R. B. Voltan, J. I. Fahl, and M. L. Carelli, Variaçao na anatomia foliar de cafeeiros submetidos a diferentes intensidades lumiosas, Rev. Bras. Fisiol. Veg, vol.4, pp.99-105, 1992.

P. G. Waterman, J. A. Ross, and D. B. Mckey, Factors affecting levels of some phenolic compounds, digestibility, and nitrogen content of the mature leaves ofBarteria fistulosa (Passifloraceae), Journal of Chemical Ecology, vol.7, issue.3, pp.387-401, 1007.
DOI : 10.1111/j.1095-8312.1981.tb01646.x

R. Zhou, W. H. Su, G. F. Zhang, Y. N. Zhang, and X. R. Guo, Relationship between flavonoids and photoprotection in shade-developed Erigeron breviscapus transferred to sunlight, Photosynthetica, vol.21, issue.2, pp.201-209, 2016.
DOI : 10.1002/rcm.3166

M. ?iv?ák, M. Bresti?, K. Ol?ovská, and P. Slamka, Performance index as a sensitive indicator of water stress in Triticum aestivum L, Plant Soil Environ, vol.54, pp.133-139, 2008.