C. A. Acosta, Impending trade suspensions of Caribbean queen conch under CITES. Fisheries, vol.2, p.31, 2006.

A. Aranda, D. Oxenford, H. A. Bissada, C. Enriquez-díaz, M. Brulé et al., Reproductive patterns of queen conch, Strombus gigas (Mollusca, Gastropoda), across the Wider Caribbean Region, Bulletin of Marine Science, vol.90, issue.3, pp.813-831, 2014.

W. Back, B. B. Hanshaw, T. E. Pyle, L. N. Plummer, and A. E. Weidie, Geochemical significance of groundwater discharge and carbonate solution to the formation of Caleta Xel Ha, Water Resources Research, vol.15, issue.6, pp.1521-1535, 1979.

P. A. Beddows, P. L. Smart, F. F. Whitaker, and S. L. Smith, Decoupled fresh-saline groundwater circulation of a coastal carbonate aquifer: Spatial patterns of temperature and specific electrical conductivity, Journal of Hydrology, vol.346, issue.1-2, pp.18-32, 2007.

C. Berg, J. Ward, B. Luckhurst, K. Nisbet, and F. Couper, Observations of breeding aggregations of the queen conch, Strombus gigas, Bermuda. Proceedings of the Gulf and Caribbean Fisheries Institute, vol.42, pp.161-171, 1992.

E. M. Boman, M. D. Graaf, L. A. Nagelkerke, A. W. Stoner, C. E. Bissada et al., Variability in size at maturity and reproductive season of queen conch Lobatus gigas (Gastropoda: Strombidae) in the Wider Caribbean Region, Fisheries Research, vol.201, pp.18-25, 2018.

G. Ceballos, P. R. Ehrlich, A. D. Barnosky, A. García, R. M. Pringle et al., Accelerated modern human -induced species losses: Entering the sixth mass extinction, Sciences Advances, issue.1, pp.1-5, 2000.

J. A. Coyer, G. Hoarau, G. A. Pearson, E. A. Serrão, W. T. Stam et al., Convergent adaptation to a marginal habitat by homoploid hybrids and polyploid ecads in the seaweed genus Fucus, Biology Letters, vol.2, issue.3, pp.405-408, 2006.

J. C. Crawford, C. K. Nielsen, and E. M. Schauber, Survival and habitat use of sympatric lagomorphs in bottomland hardwood forests, Canadian Journal of Zoology, vol.96, issue.7, pp.713-722, 2018.

D. Oficial-de-la-federación, Acuerdo por el que se modifica el similar por el cual se establecen periodos de veda para la pesca comercial de caracol rosado o blanco (Strombus gigas) en aguas de jurisdicción federal correspondientes al litoral del Estado de Quintana Roo, 2017.

J. C. Doerr and R. L. Hill, Home range, movement rates, and habitat use of Queen Conch, Strombus gigas, Caribbean Journal of Science, vol.47, issue.2, pp.251-259, 2013.

R. C. Duarte, A. A. Flores, C. Vinagre, and M. C. Leal, Habitatdependent niche partitioning between colour morphs of the algal-dwelling shrimp Hippolyte obliquimanus, Marine Biology, vol.164, issue.11, p.215, 2017.

A. M. Dujon, T. C. Stieglitz, E. Amice, and D. M. Webber, Snail leaps and bounds: Drivers of the diel movement pattern of a large invertebrate, the Caribbean queen conch (Lobatus gigas), in a marginal inshore habitat, Canadian Journal of Zoology, vol.97, pp.436-445, 2019.

S. D. Gaines, C. White, M. H. Carr, and S. R. Palumbi, Designing marine reserve networks for both conservation and fisheries management, Proceedings of the National Academy of Sciences of the United States of America, vol.107, pp.18286-18293, 2010.

R. A. Glazer and J. A. Kidney, Habitat associations of adult queen conch (Strombus gigas L.) in an unfinished Florida keys back reef: Applications to essential fish habitat, Bulletin of Marine Science, vol.75, issue.2, pp.205-224, 2004.

B. R. Gondwe, S. Lerer, S. Stisen, L. Marín, M. Rebolledo-vieyra et al., Hydrogeology of the south-eastern Yucatan Peninsula: New insights from water level measurements, geochemistry, geophysics and remote sensing, Journal of Hydrology, vol.389, issue.1-2, pp.1-17, 2010.

A. Grüss, D. M. Kaplan, S. Guénette, C. M. Roberts, and L. W. Botsford, Consequences of adult and juvenile movement for marine protected areas, Biological Conservation, vol.144, issue.2, pp.692-702, 2011.

B. S. Halpern, S. Walbridge, K. A. Selkoe, C. V. Kappel, F. Micheli et al., A global map of human impact on marine ecosystems, Science, vol.319, pp.948-953, 2008.

M. R. Heithaus and L. M. Dill, Food availability and tiger shark predation risk influence bottlenose dolphin habitat use, Ecology, vol.83, issue.2, pp.480-491, 2002.

M. R. Heithaus, A. Frid, A. J. Wirsing, L. M. Dill, J. W. Fourqurean et al., State-dependent risk-taking by green sea turtles mediates top-down effects of tiger shark intimidation in a marine ecosystem, Journal of Animal Ecology, vol.76, issue.5, pp.837-844, 2007.

K. O. Hesse, Movement and migration of the queen conch, Strombus gigas, in the Turk and Caicos Islands, Bulletin of Marine Science, vol.29, issue.3, pp.303-311, 1979.

J. L. Hopwood, The contribution of roadside grassland restorations to native bee conservation, Biological Conservation, issue.10, pp.2632-2640, 2008.

C. Huveneers, C. A. Simpfendorfer, S. Kim, J. M. Semmens, A. J. Hobday et al., The influence of environmental parameters on the performance and detection range of acoustic receivers, Methods in Ecology and Evolution, vol.7, issue.7, pp.825-835, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01986390

M. L. Imhoff, L. Bounoua, T. Ricketts, C. Loucks, R. Harriss et al., Global patterns in human consumption of net primary production, Nature, vol.429, pp.870-873, 2004.

J. B. Jackson, M. X. Kirby, W. H. Berger, K. A. Bjorndal, L. W. Botsford et al., Historical overfishing and the recent collapse of coastal ecosystems, Science, vol.293, issue.5530, pp.629-637, 2001.

?. Jankowiak and ?. ?awicki, Marginal habitats as important refugia for riparian birds during flood years, Bird Study, issue.1, pp.125-129, 2014.

T. J. Kawecki, Adaptation to marginal habitats, Annual Review of Ecology, Evolution, and Systematics, vol.39, issue.1, pp.321-342, 2008.

G. I. Kerley, R. Kowalczyk, and J. P. Cromsigt, Conservation implications of the refugee species concept and the European bison: King of the forest or refugee in a marginal habitat?, Ecography, vol.35, issue.6, pp.519-529, 2012.

A. S. Kough, C. A. Belak, C. B. Paris, A. Lundy, H. Cronin et al., Ecological spillover from a marine protected area replenishes an over-exploited population across an island chain, Conservation Science and Practice, vol.1, issue.3, 2019.

D. Lagomasino, R. M. Price, J. Herrera-silveira, F. Miralles-wilhelm, G. Merediz-alonso et al., Connecting groundwater and surface water sources in groundwater dependent coastal wetlands and estuaries: Sian Ka'an Biosphere Reserve, vol.38, pp.1744-1763, 2015.

R. J. Larson, Competition, habitat selection, and the bathymetric segregation of two rockfish (Sebastes) species, Ecological Monographs, vol.50, issue.502, pp.221-239, 1980.

P. Lozano-Álvarez and E. Briones-fourzán, Aspectos poblacionales del Caracol Rosado (Strombus gigas) y las langostas (Panulirus argus y P. guttatus) del parque Chankanaab, Biodiversidad Acuática de la Isla de Cozumel, pp.279-307, 2007.

F. J. Mazzotti, H. E. Fling, G. Merediz, M. Lazcano, C. Lasch et al., Conceptual ecological model of the Sian Ka'an biosphere reserve, vol.25, pp.980-997, 2005.

E. S. Minor and D. L. Urban, A graph-theory framework for evaluating landscape connectivity and conservation planning, Conservation Biology, vol.22, issue.2, pp.297-307, 2008.

J. R. Peel and D. Aranda, Growth, recruitment, and mortality of the Queen Conch Strombus gigas in a natural protected area of the Mexican Caribbean, Proceedings of the 65th Gulf and Caribbean Fisheries Institute, pp.446-449, 2012.

J. R. Peel and D. Aranda, Growth and population assessment of the queen conch Strombus gigas (Mesogastropoda: Strombidae) by capture mark-recapture sampling in a natural protected area of the Mexican Caribbean, Revista de Biología Tropical, vol.60, pp.127-137, 2012.

E. Perry, G. Velazquez-oliman, and L. Marin, The hydrogeochemistry of the karst aquifer System of the Northern Yucatan Peninsula, Mexico. International Geology Review, vol.44, issue.3, pp.191-221, 2002.

J. M. Posada, A. W. Stoner, K. S. Sealey, A. Antczak, R. Torres et al., Regional initiative for the evaluation of queen conch (Strombus gigas) exploitation under an historical perspective, Proceedings of the 59th Gulf Caribbean Fisheries Institute, pp.23-30, 2006.

H. R. Pulliam, R: A language and environment for statistical computing, R Foundation for Statistical Computing, vol.3, pp.349-361, 2000.

T. C. Stieglitz and A. M. Dujon, A groundwater-fed coastal inlet as habitat for the Caribbean queen conch Lobatus gigas -An acoustic telemetry and space use analysis, Marine Ecology Progress Series, vol.571, pp.139-152, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01765577

R. K. Stoessell, W. C. Ward, B. H. Ford, and J. D. Schuffert, Water chemistry and CaCO 3 dissolution in the saline part of an open-flow mixing zone, vol.101, pp.159-169, 1989.

A. W. Stoner, What constitutes essential nursery habitat for a marine species? A case study of habitat form and function for queen conch, Marine Ecology Progress Series, vol.257, pp.275-289, 2003.

A. W. Stoner, M. H. Davis, and C. J. Booker, Abundance and population structure of queen conch inside and outside a marine protected area: Repeat surveys show significant declines, Marine Ecology Progress Series, vol.460, pp.101-114, 2012.

A. W. Stoner, K. W. Mueller, N. J. Brown-peterson, M. H. Davis, and C. J. Booker, Maturation and age in queen conch (Strombus gigas): Urgent need for changes in harvest criteria, Fisheries Research, vol.131, pp.76-84, 2012.

A. W. Stoner and M. Ray-culp, Evidence for Allee effects in an over-harvested marine gastropod: Density-dependent mating and egg production, Marine Ecology Progress Series, vol.202, pp.297-302, 2000.

A. W. Stoner and K. C. Schwarte, Queen conch, Strombus gigas, reproductive stocks in the central Bahamas: Distribution and probable sources, Fishery Bulletin, vol.92, issue.1, pp.171-179, 1994.

J. A. Tello-cetina, L. A. Rodríguez-gil, and F. Rodríguez-romero, Genética poblacional del caracol rosado Strombus gigas en la Península de Yucatán: Implicaciones para su manejo y pesquería, Ciencias Marinas, vol.31, issue.2, pp.379-386, 2005.

A. Tewfik, E. A. Babcock, R. S. Appeldoorn, and J. Gibson, Declining size of adults and juvenile harvest threatens sustainability of a tropical gastropod, Lobatus gigas, fishery, Aquatic conservation: Marine and freshwater ecosystems, pp.1-21, 2019.

S. Theile, Queen conch fisheries and their management in the Caribbean. Technical report to the CITES secretariat in completion of contract, 2001.

N. K. Truelove, S. J. Box, K. A. Aiken, A. Blythe-mallett, E. M. Boman et al., Isolation by oceanic distance and spatial genetic structure in an overharvested international fishery, Diversity and Distributions, vol.23, issue.11, pp.1292-1300, 2017.

M. Valle-esquivel, Growth of queen conch, Strombus gigas, Proceedings of the Gulf and Caribbean Fisheries Institute, vol.50, pp.66-77, 1998.

S. M. Vrdoljak and R. C. Hart, Groundwater seeps as potentially important refugia for freshwater fishes on the Eastern Shores of Lake St Lucia, African Journal of Aquatic Science, vol.32, issue.2, pp.125-132, 2007.

B. A. Wintle, H. Kujala, A. Whitehead, A. Cameron, S. Veloz et al., Global synthesis of conservation studies reveals the importance of small habitat patches for biodiversity, Proceedings of the National Academy of Sciences of the United States of America, vol.116, pp.909-914, 2019.

M. Zato?, O. Vinn, and A. M. Tomescu, Invasion of freshwater and variable marginal marine habitats by microconchid tubeworms -an evolutionary perspective, Geobios, vol.45, issue.6, pp.603-610, 2012.