Mediterranean wetlands are of key interest in the preservation of biodiversity. However, their ecological water requirement is in constant competition with human uses, particularly in a context of aridity and climate change. This paper examines the case of the Ichkeul Lake-Lagoon system located in the north of Tunisia in a river basin that is heavily contributing to the water supply of southern regions through water transfers. It maps out the various actors involved, their stakes, discourses and strategies, and explains the current degradation of the Ichkeul system by unpacking the decision-making power of these actors. It confirms that marginal social groups, ecosystems and next generations are the main parties affected by current water allocation patterns.